This paper describes the osteological features in the context of the structural complexes they build as a way of avoiding their separation from the morphofunctional wholes to which they belong. The posterior zone of the neurocranium is compact, and the splanchnocranium is laminar with very little cartilaginous tissue sustaining robust oral jaws. The pharyngobranchial apparatus is short anteroposteriorly and expanded laterally.The hyoid bar is small, about as long as its branchiostegal rays. The spatial relationship of the paired and unpaired fins to the vertebral column was highly constant in the specimens examined of Helcogrammoides chilensis from the Central zone of Chile. The absence of a neural spine in the first two vertebrae, a highly fused caudal complex, the urohyal with well developed lateral processes, and presence of a pelvic spine, are osteological features that contribute to the diagnosis of Helcogrammoides.